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Inquisitor 1391

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Ripper | 11:25 Sun 21st Jun 2015 | Crosswords
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Can anyone help with 43 down? (there may be an item of headwear missing)
A selection of English rodents (4) G?IS
Also I don't understand the preamble about what to highlight "the section of the grid in which the relevant callers are located" any hints would be appreciated!
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Glis, rodents - part of 'English'.
Once you've found the first caller, the rest will follow - literally. The title might help.
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Thank you Neveracrossword enjoy the sun!

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